Zonation On Rocky Shore Essay, Research Paper
The seashore is a
habitat that contains a wide range of microhabitats and ecological niches for
different creatures. This is mainly due to the effects of the tides, that rise
and fall twice each day. Tides are the vertical movement of water in a
periodical oscillation of the sea, due to the gravitational pull of the sun and
moon. The tides are on a semi-diurnal cycle, so there are two high tides and two
low tides each day. Due to the orbit of the moon, the tides also have a monthly
cycle. This creates neap (very low) and spring (very high) tides. The seashore
can be divided into several zones, which are illustrated on the diagram below:
Key: EHWS = Extreme High Water Spring (MHWS = Mean High Water Spring) MHWN =
Mean High Water Neap (MTL = Mid Tide Level) MLWN = Mean Low Water Neap ELWS =
Extreme Low Water Spring (MLWS = Mean Low Water Spring) CD = Chart datum The
Supralittoral Zone: This is the highest zone on the shore, and lies above the
EHWS mark, and therefore is never covered by seawater. However, it may be
occasionally be spray wetted. Because of this, it is mainly inhabited by
terrestrial species, such as lichen, that can live in areas of very high
salinity. The Littoral (Intertidal) Zone: This zone is the area that is covered
and uncovered by the tides, and therefore organisms that live here must be able
to tolerate a large range of conditions. It can be further divided into the
Littoral Fringe and the Eulittoral zone. The Littoral Fringe (Splash Zone): This
part of the Littoral zone lies above the area that is completely submerged by
the sea in normal conditions. However, it is frequently covered by splash from
waves, and so is far more marine in character that the Supralittoral Zone.
Lichens still dominate this zone, but some species of periwinkles and topshells
may graze them. The Eulittoral Zone: This zone is the area of the beach that is
regularly submerged by the tides, and can be divided into three more zones, the
upper, middle and lower shores. It shows the greatest species diversity of any
of the zones. The Upper Shore: This region of the shore lies between the EHWS
and MHWN marks, and so is only immersed during spring tides. Because of this,
organisms that live here must be adapted to survive long periods of desiccation.
The two seaweeds that are the most common here, Fucus spiralis and Pelvetia
canaliculata have adaptations to survive in this area. The Middle Shore: This
region of the shore lies between the MHWN and MLWN marks, and will be submerged
for half of every day, even during neap tides. The most common seaweed in this
zone Fucus vesiculosus. Mussel beds will form and both limpets and periwinkles
will graze the rocks. Sea anemones and crabs are residents of this zone. The
Lower Shore: This region of the shore lies between the MLWN and ELWS marks, and
will be submerged for most of each day, even during neap tides. The most
important seaweed in this area is Fucus serratus, which will form large zones
wherever suitable. It shows the greatest species diversity of any zone on the
seashore. The Sublittoral Zone: This part of the shore lies below the ELWS mark,
and is therefore never uncovered by the sea. There are many types of organism
found on the rocky shore. The two main photosynthetic organisms are the lichens
and the macroalgae or seaweeds. Lichen are the main organisms found in the
splash zone and come in three distinct types; crustose, foliose and fruiticose.
Crustose lichens form a thin crust on the rock surface, and are impossible to
remove without damage. Foliose lichens are leafy lichens that are not as firmly
attached to the rocks. Fruiticose lichens extent vertically from the rock
surface, and can sometimes be confused with mosses and small grasses. The leafy
part of a lichen is known as the thallus. Seaweeds are primarily divided by
colour, into brown, red and green groups. Most marine seaweeds are brown
seaweeds, with fewer red species, and even fewer green species. The three main
parts of a seaweed are: 1. Frond (lamina, thallus, blade) (often broad and flat)
2. Stipe region (often long and cylindrical) 3. Basal attachment (holdfast) The
frond or thallus is the site of most of the photosynthetic activity in the
organism, and also contains the reproductive organs. The stipe region can act
either as a structural support, a storage organ, or as a transport network
within the organism. The role of the holdfast is to anchor the seaweed securely
to the substrate it lives on. The holdfast must be strong enough to resist the
strong pull of the waves and tides on the seaweed. The size and strength of the
holdfast varies between species. The main heterotrophic organisms of the
seashore are the molluscs. The most common molluscs are the gastropods
(periwinkles, limpets and topshells), and the mussels. Periwinkles have coiled
shells and a circular operculum (a small, retractable piece of shell used to
cover the opening of the shell when the snail is inside.). They average about
15mm in length and are the most common group of gastropods on the seashore.
Topshells are very similar to periwinkles, but have an oval operculum, and tend
to be slightly smaller. There are fewer species of topshells than periwinkles on
a rocky shore. Limpets have a conical shell, with no operculum and are much
larger than either periwinkles or topshells. Mussels have two shells, and are
fixed to a single location in adult life. They can form large groups on the
rocky shore. Describe LOWER SHORE There was only one species of seaweed found in
the lower shore, Fucus serratus, and it was very abundant. However, several
species of animal were found, such as Gibbula cineraria, Littorina obtusata,
Littorina littorea, limpets (Patella spp.) and mussels (Myttilus edulis). Of
those, Gibbula cineraria was the most abundant. Fucus serratus: This species of
brown seaweed (Phaoephyta) was found only below the MLWN mark in stations 10, 11
and 12. It was most common in station 11 (40% cover), but there was not a lot of
difference in the distributions between these three stations. Fucus serratus is
a medium sized marine seaweed with a flattened, branched thallus with a small
stipe for support and a small holdfast. At the ends of the thalli, there are
small, swollen areas called receptacles, which contain many conceptacles, in
which gamete production occurs. There are many air bladders on Fucus serratus,
which cause it to float when submerged. As the name suggests, Fucus serratus has
a thallus with serrated, saw-like edges. Gibbula cineraria: This species of
topshell was found mainly in the lower shore, below the MLWN mark (stations
10,11, and 12), and in station 9 (just above the MLWN mark). It was evenly
distributed across stations 9, 10, and 11, with similar numbers in each quadrat
(between 40 and 50 individuals per quadrat). It was far less common in station
12, where only two individuals were found. Gibbula cineraria is a relatively
large snail, at just over 15-mm. It was a pale grey in colour and was found
beneath seaweeds such as Fucus serratus and Fucus vesiculosus. MIDDLE SHORE
Several species of seaweed were recorded in the middle shore. Fucus vesiculosus,
Ascophyllum nodosum and Polysiphona lanosa were all found, and Fucus vesiculosus
was the most abundant. Many animal species were recorded, such as Gibbula
umbilicalis, G. cineraria, Littorina saxatalis, L. obtusata, L. littorea,
limpets (Patella spp.) and mussels (Myttilus edulis). Of these Gibbula cineraria
was the most abundant. Fucus vesiculosus: This seaweed was found mainly in the
middle shore, between the MLWN and MHWN marks (stations 7,8 and 9), but also in
station 6 (just above the MHWN mark). There was a much lower density in stations
6,7 and 8 (between 3 and 12%), than in station 9, where the percentage cover was
30%. Fucus vesiculosus is similar to Fucus serratus (see above), with a
flattened, branched thallus and air bladders, but lacks the serrated edges of
Fucus serratus. Littorina obtusata agg.: This species of periwinkle was found in
the middle shore (stations 7,8 and 9) and the lower upper shore (station 6). It
was also recorded in station 12, at the lower end of the lower shore. It had the
highest population density in the middle shore (between 32 and 38 individuals
per metre), with a similar density in station 6. It was far less abundant in
station 12, with only 12 individuals recorded. Littorina obtusata agg. is a
small, flat periwinkle, mainly found on the underside of seaweeds such as Fucus
vesiculosus, Fucus spiralis and Ascophyllum nodosum, where it mimics air
bladders. It comes in a wide range of colour, but most individuals are a dark
olive green to match the seaweeds they live on. UPPER SHORE Again, several
species of seaweed were recorded in this zone, such as Fucus vesiculosus, F.
spiralis, Ascophyllum nodosum, Pelvetia canaliculata and Polysiphona lanosa.
Several animal species were also recorded, such as Littorina saxatalis, L.
obtusata and limpets (Patella spp.) Pelvetia canaliculata: This seaweed was
found in station 4 only (at the very upper limit of the littoral zone, just
below the EHWS mark), but was very abundant, covering 70% of the quadrat.
Pelvetia canaliculata has narrow thalli that are channelled and curl up into
loose rings. It is browny red in colour and has no air bladders for support.
Littorina saxatalis: This species of periwinkle was found across the whole upper
shore (stations 4,5 and 6) and at the top of the middle shore (station 7). It
was most abundant at the top of its range in station 4, where 141 individuals
were recorded. It became less and less abundant down the beach, at the bottom of
its range, in station 7, where only 20 individuals were recorded. Littorina
saxatalis is a medium-sized periwinkle, about 16-mm long. It has a ridged shell
that is orange-brown in colour, and is commonly found in crevices and cracks on
the upper shore. SPLASH ZONE The only plants found in the splash zone where
lichens such as Verrucaria maura, Xanthoria parientina, Ramalina siliquosa,
Lecanora atra and Ochrolechia parella. No animal species were recorded in this
zone. Xanthoria parientina: This species of foliose lichen was found throughout
the splash zone (stations 1,2 and 3), and was the largest range out of all the
lichens. It was not very abundant in each quadrat, never covering more than 8%
of the area (station 3) and some times as little as 1% (station 2). Xanthoria
parientina is a foliose lichen, which means it is only loosely attached to the
rock, and has large thalli. It was orangey yellow in colour. Explain The
environmental gradient on the seashore is constantly changing. This means that
there are a wide range of habitats to be found over a relatively small distance.
The wide range of species found on the seashore is due to the wide range of
habitats and conditions found there. Species can only be adapted to a small
range of conditions, so as the conditions on the seashore change, so do the
species found there. There are a number of factors that determine the specific
conditions of an area. These factors can be either biotic or abiotic. Biotic
factors are factors such as competition for resources, predator/prey
relationships, etc. Abiotic factors are factors like temperature, relief,
climate, etc. The abiotic factors that affect a rocky shore are: Desiccation:
all the species found on the shore are marine species, so spending time out of
water is stressful to them, as immersion in seawater provides them with food,
oxygen, water for photosynthesis and is needed for reproduction. Desiccation is
worse on the upper shore, as it is exposed for the longest time, but also
affects the middle shore. Temperature: Seawater remains at a far more constant
temperature that the land, (seawater varies between 5? and 15? Celsius,
whereas the land temperature varies between below freezing in winter and 30? C
plus in summer) so species that are immersed in seawater for long periods of
time are buffered against large temperature changes. The temperature of the
surroundings also affects the rate of metabolism; very cold conditions will slow
it down, whereas very high temperatures may denature vital enzymes. Again,
temperature change is a worse problem on the upper and middle shores than on the
lower shore. Wave action: The action of powerful waves can dislodge many
species, so those that live on the middle shore (where wave action is at its
most powerful) must be adapted to survive very rough conditions. Wave action
also increases the humidity of an area, and so can help to reduce desiccation.
Light: Light is needed for photosynthesis, and all seaweeds must be immersed in
water for this to occur. Water filters off some of the wavelengths of light and
reduces the intensity that reaches the seaweeds. To maximise the light that does
reach them red and brown seaweeds have accessory pigments that help to absorb
different wavelengths of light. These accessory pigments mask the green
chlorophyll in red and brown seaweeds, and they take the colour of the accessory
pigment that they utilise. Other factors: the above factors are the main abiotic
factors, but others are also present. The aspect of a slope affects the
temperature and rate at which water evaporates, so south facing slopes are
warmer, but dry faster, while north facing slopes are cooler and damper. The
steepness of a slope also affects the rate at which it drains, as a steeper
slope drains faster than a shallower one, so desiccation is more of a problem.
The turbidity or cloudiness of seawater (due to plankton, sewage and other
detritus) can affect the intensity of light reaching submerged seaweeds. Another
factor is the seepage of freshwater onto the shore. Many seaweeds cannot
tolerate salinity changes, so other species that can tolerate such changes will
inhabit these areas. The biotic factors that affect the rocky shore tend to
affect the lower limits at which a species may live. The biotic factors that
affect the distribution of organisms on the rocky shore are: Food supply: All
organisms need food to survive and so can only flourish in areas in which they
can find food. Many species that are found on the seashore left the sea in
search of food supplies. For organisms, such as barnacles, which depend on food
carried by the waves, far more food will be found in the intertidal zone that at
the bottom of the sea. Predation: Many species also live on the seashore in an
attempt to evade marine predators, such as fish, crabs, lobsters etc, that are
far more common in the sea than on the shore. Organisms will also try to live as
far up the shore as possible in order to avoid their less well adapted
predators. Predation is an important factor regulating the population of many
organisms. Reproduction: Most marine organisms still rely on the sea for
reproduction, so animal species, such as crabs, may migrate lower down the shore
in order to release their gametes. Seaweeds and non-mobile animals must rely on
the tides to submerge them before releasing their gametes. Competition: This is
the most important biotic factor determining the distribution of species on the
seashore. There are two types of competition, interspecific (between two
different species) and intraspecific (between individuals of the same species).
Organisms compete for all the resources that are in short supply. On the
seashore, most resources are in short supply, so organisms compete for space,
food, and light. Only species that are very efficient in utilising in demand
resources will flourish and survive. Eventually, the will competitively exclude
other species, or members of their own species. Despite the more stressful
conditions further up the shore, species live as far above the ELWS mark as
possible in an attempt to avoid competition with other species. For example,
Fucus spiralis is very well adapted to surviving long periods out of water, so
it is found in the upper shore. It is not found in the middle and lower shores
because competition with other species of seaweeds such as Fucus vesiculosus and
Fucus serratus prevents them from surviving, so no specimens are found. Species
can adapt to these different factors in three ways. They can adapt in physical,
physiological or behavioural ways. Physical adaptations are those that modify
the external appearance of an organism, physiological adaptations are those that
modify the internal organisation of an organism and behavioural adaptations are